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Harrismith Nature Reserve

25/11/2021

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​Greetings fellow foxies,
Harrismith reserve adjoins the township of Harrismith, 73 kilometres east of Narrogin and 55 kilometres south-east of Wickepin. . The Oasis Hotel has accommodation, lunches, drinks and snacks. With an adjoining caravan park, Harrismith is a good spot to camp overnight and enjoy the wildflowers. 
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The reserve differs from others closer to Narrogin in having little woodland. It is on a broad ridge near the divide between the Swan-Avon and Blackwood River catchments.
The landscape is a lateritic gravel plain that formed millions of years ago when the climate was wetter. Unusual circular hollows were once wetlands overlying granite base-rock.
As the climate became drier, wetlands dried out and were colonised by eucalypt and sheoak trees. Lateritic heath persisted on the surrounding gravelly plain and built up a raised  doughnut shaped edge around each hollow.
Soils are mainly shallow lateritic ironstone and gravels with deeper white sandy topsoils in broad depressions. They are infertile and have poor water retention.  
It is first class kwongan heath plain. Many heath plants flower after woodland wildflowers in other reserves have finished.
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The 3.5 kilometre gently sloping trail (level 2) is an easy walk, or cycle using a mountain/comfort bike. There is also a drive trail. The airport section has particularly good wildflowers but keep to the side as it is a working airstrip.
The best time to visit is from mid-August to November, with October being peak time for verticordias and other flowering shrubs. As there are few trees on this walk take water and walk at cooler times on hot days.
Ignore the small redundant sign at the trailhead that indicates three walk trails and follow the blue arrows/tall red-banded posts. Orchids flower in early/mid September, but many kwongan plants flowers later. At least 2 trips are needed to see the full wildflower spectrum.
This document contains a pictorial list of wildflowers in the reserve.
harrismith_wildflowers_comp.pdf
File Size: 3665 kb
File Type: pdf
Download File

​Numbers on the diagram indicate described trail sections
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Numbers indicate sections of the trail described below
Section 1 is a gently sloping gravelly track through medium to tall kwongan shrubland, which is interspersed by small rock sheoak and cypress pine trees. Check for jug orchids (Pterostylis recurva) and green spider orchids (Caladenia falcata) in September.
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1 Mixed level shrubland
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1 Trail in the early morning
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1 Verticordia serrata at start of trail
Section 2 starts at the picnic table where shrubland opens out at a very broad yellow sand hollow containing a few groves of eastern wandoo (Eucalyptus capillosa) trees). I found a few stark white spider orchids (Caladenia longicauda subsp. eminems) in early September Beautiful Verticordias picta, roei, and serrata flower in late September to October.
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2 Verticordia picta and Calytrix lechenaultii
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2 Picnic table
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2 Verticordia roei, Hakea multilineata trees
An unmarked trail goes east from the picnic table to open wandoo and rock sheoak woodland (Section 3). Chameleon spider orchids (Caladenia dimidia), sugar orchids (Caladenia saccharata), common donkey orchids (Diuris corymbosa) and cowslip orchids (Caladenia flava) flower here in early to mid-September. Return to the main track to avoid getting lost, because the side trail continues for a long way before becoming a dead end. 
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3 Diuris corymbosa early September
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3 Waitzia acuminata October
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3 Caladenia dimidia early September
The trail continues to a slight grey sandy slope (section 4) containing sphere banksia (Banksia sphaerocarpa), Lambertia ilicifolia), and roadside tea trees (Leptospermum erubescens) interpersed by rock sheoak and spindly mallees. Check the ground for cowslip orchids and painted sundew (Drosera zonaria). 
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4 Lambertia ilicifolia in September
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Sandy trail on section 4
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4 Crowded Drosera zonaria in September
Picture5 Cowslip/little pink fairy orchid hybrid
At section 5 the trail passes through prickly gravel scrub. This is the rim of 'doughnut' valley to the south, where you can see tree branches.
​There is no trail down to the valley, but a moderately easy 20 metre scramble down the slope reveals attractive woodland. There is a variety of everlastings and other flowers on the slope and valley. if you are lucky, you may find the cowslip/little pink fairy orchid hybrid.
​
Cars can reach the picnic table on the trail adjoining the airstrip by using the drive trail. 

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5 Trail along valley rim
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5 Phebalium tuberculosum late September
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5 Rhodanthe manglesii in valley
PictureSouthern view along airstrip in mid October
​Section 6 follows the airstrip. You can see sheet ironstone at about 40cm below the surface. It is amazing that such a variety of of plants grow here. large shrubs flourish because they have taproots, which penetrate cracks in the ironstone.
Vegetation on the uphill side of the airstrip is larger because of a chemical overspray, which killed large shrubs on the downhill side. Surprisingly there appears to be little long-term damage as smaller shrubs have already compensated. However, some kwongan plants depend on fire for seed germination and may not regenerate until this occurs.
There is a wonderful show of flowering shrubs by the airstrip in late September and October. Walk down the eastern airstrip to meet the track and return.


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Dasymalla terminalis October
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Grevillea eryngioides late October
Conclusion
Harrismith reserve is a great spot for wildflower enthusiasts in spring. The trail and Toolibin Nature Reserve would make a good day trip from Narrogin or Wickepin.
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Inside a Sun Orchid

14/11/2021

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Greetings fellow Foxies,
I have been into Thelymitra/sun orchid flowers lately.
Sun orchids have normal petals and sepals like many other flowers. When a Foxes Lair visitor found it difficult to believe that they were orchids, I couldn’t give a simple answer why. It is the fused pistil-stamen structure called the column, but simple it is not, and discovering why was a real journey for me!
The image below shows male (stamens) and female (pistil) parts of a fairly typical Ixia lily.
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Section of a non-orchid flower showing male and female flower parts
​In our orchids, the filaments and style have been fused to form the column, which usually contains anthers at the top and the stigma at the base (reduced to a stigmatic plate).
Most orchids aso have a modified bottom petal called a labellum, which acts as a landing ramp and an insect attractor.
​Pollinators are led up the labellum to the stigmatic plate. In doing so they unknowingly collect pollen by rubbing against the anthers at the top of the column, and drop pollen that they accumulated from another orchid.
Blue,cowslip, and sun orchids do not suppy pollen or nectar for pollinators. They rely on visual deception by resembling flowers such as lilies and hibbertias  which do.
One would think it would be easier to supply a bit of nutrition. Perhaps the energy saved from being stingy gives them an edge in very dry seasons. Some produce a scent, most don’t. 
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Orthroxanthus laxus lily
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Scented sun orchid T.macrophylla
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Blue china orchid
​Sun orchids lack a labellum, and their anthers are not at the top of the column. 
After many hours pondering over the structure of a sun orchid column, I realised that the shape is a compensation for the lack of a labellum, and designed to entice insects to land directly on the column.
The column has an oval sheath below (wing), which surrounds the anthers and stigmatic plate. It pinches at the top and opens again as a smaller hood, to give the overall shape of a hooded cloak.  Thelymitra is derived from Greek words for ‘woman’s hood’
The hood is the most distinctive part of the column, often shaped and coloured like an anther, and having knobs, wings,’ hairs’ and sometimes scent to attract insects. I thought the yellow hood was an anther. It certainly fooled me as well as the hoverfly below (which was too large to pollinate the orchid).
In the wing cavity anthers are separated from the stigmatic plate by a pouch, with a knob on top called the rostrum. 
 How does it operate? Small pollinating bees would land in the hood area, then tip backward to contact anthers underneath and drop other pollen on to the stigmatic plate.
​
​Apparently, the anthers from insect pollinated plants remain intact and stick to the bee as a unit. Anthers on self-pollinated species break up, allowing pollen to fall down to the stigma plate.
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Scented sun orchid column
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Text book sun orchid column description
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Hover fly probing sun orchid hood, Image Lyn Alcock
With flower colour, the shape and protuberences of the hood and wing distinguish sun orchid species. See the variations!

The leopard orchid is the weirdest. In addition to knobs, fringes and cilia, it has a long projection (resembing a priapic little man) called an anther point, which joins the anthers . Leopard orchids are self pollinated, so perhaps bees jiggling the anther point breaks up the anthers to release pollen.The closely related York sun orchid  has a similar column. Interesting that the Lasioglossum native bee in the image is attracted to the top of the column
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Leopard orchid T.benthamiana
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York sun orchid T.yorkensis image A. Watts
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Custard orchid T.villosa
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Wandoo sun orchid T.latiloba
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Lemon scented sun orchid T. antennifera
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Blue lady orchid T. crinita
PictureWasp trying to mate with a beard orchid. Image Lyn Alcock.
Being a commited Darwinian I find it difficult to see how all these modifications benefit sun orchids when they are pollinated by similar native bees.
​
Another case in point is the beard orchid, which uses pheromones and a bearded labellum that looks like a female flower wasp to a male. Why bother when the orchid pollinates itself? Perhaps the wasp's frantic probing breaks up anthers, but it seems a bit over the top. Still, that is why nature is so fascinating.
​
 To quote Charles Darwin. 'Evolution doesn't have to be perfect, just good enough'..

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    Doug Sawkins is a friend of Foxes Lair 

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